Big Brain

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Big Brain

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The frontal lobe is associated with executive functions including self-control , planning , reasoning , and abstract thought , while the occipital lobe is dedicated to vision.

Within each lobe, cortical areas are associated with specific functions, such as the sensory , motor and association regions. Although the left and right hemispheres are broadly similar in shape and function, some functions are associated with one side , such as language in the left and visual-spatial ability in the right.

The hemispheres are connected by commissural nerve tracts , the largest being the corpus callosum.

The cerebrum is connected by the brainstem to the spinal cord. The brainstem consists of the midbrain , the pons , and the medulla oblongata. The cerebellum is connected to the brainstem by pairs of tracts.

Within the cerebrum is the ventricular system , consisting of four interconnected ventricles in which cerebrospinal fluid is produced and circulated.

Underneath the cerebral cortex are several important structures, including the thalamus , the epithalamus , the pineal gland , the hypothalamus , the pituitary gland , and the subthalamus ; the limbic structures , including the amygdala and the hippocampus ; the claustrum , the various nuclei of the basal ganglia ; the basal forebrain structures, and the three circumventricular organs.

The cells of the brain include neurons and supportive glial cells. There are more than 86 billion neurons in the brain, and a more or less equal number of other cells.

Brain activity is made possible by the interconnections of neurons and their release of neurotransmitters in response to nerve impulses.

Neurons connect to form neural pathways , neural circuits , and elaborate network systems. The whole circuitry is driven by the process of neurotransmission.

The brain is protected by the skull , suspended in cerebrospinal fluid , and isolated from the bloodstream by the blood—brain barrier.

However, the brain is still susceptible to damage , disease , and infection. Damage can be caused by trauma , or a loss of blood supply known as a stroke.

The brain is susceptible to degenerative disorders , such as Parkinson's disease , dementias including Alzheimer's disease , and multiple sclerosis.

Psychiatric conditions , including schizophrenia and clinical depression , are thought to be associated with brain dysfunctions.

The brain can also be the site of tumours , both benign and malignant ; these mostly originate from other sites in the body.

The study of the anatomy of the brain is neuroanatomy , while the study of its function is neuroscience.

Numerous techniques are used to study the brain. Specimens from other animals, which may be examined microscopically , have traditionally provided much information.

Medical imaging technologies such as functional neuroimaging , and electroencephalography EEG recordings are important in studying the brain.

The medical history of people with brain injury has provided insight into the function of each part of the brain. Brain research has evolved over time, with philosophical, experimental, and theoretical phases.

An emerging phase may be to simulate brain activity. In culture, the philosophy of mind has for centuries attempted to address the question of the nature of consciousness and the mind-body problem.

The pseudoscience of phrenology attempted to localise personality attributes to regions of the cortex in the 19th century.

In science fiction, brain transplants are imagined in tales such as the Donovan's Brain. The adult human brain weighs on average about 1.

The cerebrum , consisting of the cerebral hemispheres , forms the largest part of the brain and overlies the other brain structures.

Each hemisphere is divided into four main lobes — the frontal lobe , parietal lobe , temporal lobe , and occipital lobe. The brainstem , resembling a stalk, attaches to and leaves the cerebrum at the start of the midbrain area.

The brainstem includes the midbrain, the pons , and the medulla oblongata. Behind the brainstem is the cerebellum Latin : little brain.

The cerebrum, brainstem, cerebellum, and spinal cord are covered by three membranes called meninges. The membranes are the tough dura mater ; the middle arachnoid mater and the more delicate inner pia mater.

Between the arachnoid mater and the pia mater is the subarachnoid space and subarachnoid cisterns , which contain the cerebrospinal fluid.

The cerebrum is the largest part of the brain, and is divided into nearly symmetrical left and right hemispheres by a deep groove, the longitudinal fissure.

There are many small variations in the secondary and tertiary folds. The outer part of the cerebrum is the cerebral cortex , made up of grey matter arranged in layers.

It is 2 to 4 millimetres 0. The largest part of the cerebral cortex is the neocortex , which has six neuronal layers.

The rest of the cortex is of allocortex , which has three or four layers. The cortex is mapped by divisions into about fifty different functional areas known as Brodmann's areas.

These areas are distinctly different when seen under a microscope. The primary sensory areas receive signals from the sensory nerves and tracts by way of relay nuclei in the thalamus.

Primary sensory areas include the visual cortex of the occipital lobe , the auditory cortex in parts of the temporal lobe and insular cortex , and the somatosensory cortex in the parietal lobe.

The remaining parts of the cortex, are called the association areas. These areas receive input from the sensory areas and lower parts of the brain and are involved in the complex cognitive processes of perception , thought , and decision-making.

The temporal lobe controls auditory and visual memories , language , and some hearing and speech.

The cerebrum contains the ventricles where the cerebrospinal fluid is produced and circulated. Below the corpus callosum is the septum pellucidum , a membrane that separates the lateral ventricles.

Beneath the lateral ventricles is the thalamus and to the front and below this is the hypothalamus. The hypothalamus leads on to the pituitary gland.

At the back of the thalamus is the brainstem. The basal ganglia , also called basal nuclei, are a set of structures deep within the hemispheres involved in behaviour and movement regulation.

The ventral striatum consists of the nucleus accumbens and the olfactory tubercle whereas the dorsal striatum consists of the caudate nucleus and the putamen.

The putamen and the globus pallidus lie separated from the lateral ventricles and thalamus by the internal capsule , whereas the caudate nucleus stretches around and abuts the lateral ventricles on their outer sides.

Below and in front of the striatum are a number of basal forebrain structures. These include the nucleus basalis , diagonal band of Broca , substantia innominata , and the medial septal nucleus.

These structures are important in producing the neurotransmitter , acetylcholine , which is then distributed widely throughout the brain. The basal forebrain, in particular the nucleus basalis, is considered to be the major cholinergic output of the central nervous system to the striatum and neocortex.

The cerebellum is divided into an anterior lobe , a posterior lobe , and the flocculonodular lobe. It is connected to the midbrain of the brainstem by the superior cerebellar peduncles , to the pons by the middle cerebellar peduncles , and to the medulla by the inferior cerebellar peduncles.

The brainstem lies beneath the cerebrum and consists of the midbrain , pons and medulla. It lies in the back part of the skull , resting on the part of the base known as the clivus , and ends at the foramen magnum , a large opening in the occipital bone.

The brainstem continues below this as the spinal cord , [40] protected by the vertebral column.

Ten of the twelve pairs of cranial nerves [a] emerge directly from the brainstem. The human brain is primarily composed of neurons , glial cells , neural stem cells , and blood vessels.

Types of neuron include interneurons , pyramidal cells including Betz cells , motor neurons upper and lower motor neurons , and cerebellar Purkinje cells.

Betz cells are the largest cells by size of cell body in the nervous system. Types of glial cell are astrocytes including Bergmann glia , oligodendrocytes , ependymal cells including tanycytes , radial glial cells , microglia , and a subtype of oligodendrocyte progenitor cells.

Astrocytes are the largest of the glial cells. They are stellate cells with many processes radiating from their cell bodies. Some of these processes end as perivascular end-feet on capillary walls.

Mast cells are white blood cells that interact in the neuroimmune system in the brain. Some genes are shown to be brain-specific.

In all neurons, ELAVL3 is expressed, and in pyramidal neurons, NRGN and REEP2 are also expressed. GAD1 — essential for the biosynthesis of the neurotransmitter GABA — is expressed in interneurons.

Proteins expressed in glial cells include astrocyte markers GFAP and SB whereas myelin basic protein and the transcription factor OLIG2 are expressed in oligodendrocytes.

Cerebrospinal fluid is a clear, colourless transcellular fluid that circulates around the brain in the subarachnoid space , in the ventricular system , and in the central canal of the spinal cord.

It also fills some gaps in the subarachnoid space, known as subarachnoid cisterns. From here, cerebrospinal fluid circulates around the brain and spinal cord in the subarachnoid space, between the arachnoid mater and pia mater.

It is constantly being regenerated and absorbed, and is replaced about once every 5—6 hours. A glymphatic system has been described [53] [54] [55] as the lymphatic drainage system of the brain.

The brain-wide glymphatic pathway includes drainage routes from the cerebrospinal fluid, and from the meningeal lymphatic vessels that are associated with the dural sinuses, and run alongside the cerebral blood vessels.

The internal carotid arteries supply oxygenated blood to the front of the brain and the vertebral arteries supply blood to the back of the brain.

The internal carotid arteries are branches of the common carotid arteries. They enter the cranium through the carotid canal , travel through the cavernous sinus and enter the subarachnoid space.

These branches travel forward and then upward along the longitudinal fissure , and supply the front and midline parts of the brain.

They travel sideways along the sphenoid bone of the eye socket , then upwards through the insula cortex , where final branches arise. The middle cerebral arteries send branches along their length.

The vertebral arteries emerge as branches of the left and right subclavian arteries. They travel upward through transverse foramina which are spaces in the cervical vertebrae.

Each side enters the cranial cavity through the foramen magnum along the corresponding side of the medulla.

The vertebral arteries join in front of the middle part of the medulla to form the larger basilar artery , which sends multiple branches to supply the medulla and pons, and the two other anterior and superior cerebellar branches.

These travel outwards, around the superior cerebellar peduncles, and along the top of the cerebellar tentorium, where it sends branches to supply the temporal and occipital lobes.

Cerebral veins drain deoxygenated blood from the brain. The brain has two main networks of veins : an exterior or superficial network , on the surface of the cerebrum that has three branches, and an interior network.

These two networks communicate via anastomosing joining veins. Blood from the medulla and pons of the brainstem have a variable pattern of drainage, either into the spinal veins or into adjacent cerebral veins.

The blood in the deep part of the brain drains, through a venous plexus into the cavernous sinus at the front, and the superior and inferior petrosal sinuses at the sides, and the inferior sagittal sinus at the back.

Blood from here joins with blood from the straight sinus at the confluence of sinuses. Blood from here drains into the left and right transverse sinuses.

The sigmoid drains into the large internal jugular veins. The larger arteries throughout the brain supply blood to smaller capillaries.

These smallest of blood vessels in the brain, are lined with cells joined by tight junctions and so fluids do not seep in or leak out to the same degree as they do in other capillaries; this creates the blood—brain barrier.

At the beginning of the third week of development , the embryonic ectoderm forms a thickened strip called the neural plate.

These swellings are known as the primary brain vesicles and represent the beginnings of the forebrain prosencephalon , midbrain mesencephalon , and hindbrain rhombencephalon.

Neural crest cells derived from the ectoderm populate the lateral edges of the plate at the neural folds.

In the fourth week—during the neurulation stage —the neural folds close to form the neural tube , bringing together the neural crest cells at the neural crest.

Cells detach from the crest and migrate in a craniocaudal head to tail wave inside the tube. The tube flexes as it grows, forming the crescent-shaped cerebral hemispheres at the head.

The cerebral hemispheres first appear on day These areas are formed as swellings known as the three primary brain vesicles.

In the fifth week of development five secondary brain vesicles have formed. The telencephalon gives rise to the cerebral cortex, basal ganglia, and related structures.

The diencephalon gives rise to the thalamus and hypothalamus. The hindbrain also splits into two areas — the metencephalon and the myelencephalon. The metencephalon gives rise to the cerebellum and pons.

The myelencephalon gives rise to the medulla oblongata. A characteristic of the brain is the cortical folding known as gyrification. For just over five months of prenatal development the cortex is smooth.

By the gestational age of 24 weeks, the wrinkled morphology showing the fissures that begin to mark out the lobes of the brain is evident.

The first groove to appear in the fourth month is the lateral cerebral fossa. This covers the fossa and turns it into a much deeper ridge known as the lateral sulcus and this marks out the temporal lobe.

The frontal lobe is involved in reasoning, motor control, emotion, and language. The corticospinal tract carries movements from the brain, through the spinal cord , to the torso and limbs.

Gross movement — such as locomotion and the movement of arms and legs — is generated in the motor cortex , divided into three parts: the primary motor cortex , found in the precentral gyrus and has sections dedicated to the movement of different body parts.

These movements are supported and regulated by two other areas, lying anterior to the primary motor cortex: the premotor area and the supplementary motor area.

These then travel down the spinal cord , with most connecting to interneurons , in turn connecting to lower motor neurons within the grey matter that then transmit the impulse to move to muscles themselves.

The sensory nervous system is involved with the reception and processing of sensory information. This information is received through the cranial nerves, through tracts in the spinal cord, and directly at centres of the brain exposed to the blood.

Mixed motor and sensory signals are also integrated. From the skin, the brain receives information about fine touch , pressure , pain , vibration and temperature.

From the joints, the brain receives information about joint position. Sensation collected by a sensory receptor on the skin is changed to a nerve signal, that is passed up a series of neurons through tracts in the spinal cord.

The dorsal column—medial lemniscus pathway contains information about fine touch, vibration and position of joints.

The pathway fibers travel up the back part of the spinal cord to the back part of the medulla, where they connect with second-order neurons that immediately send fibers across the midline.

These fibers then travel upwards into the ventrobasal complex in the thalamus where they connect with third-order neurons which send fibers up to the sensory cortex.

The pathway fibers travel up the spinal cord and connect with second-order neurons in the reticular formation of the brainstem for pain and temperature, and also terminate at the ventrobasal complex of the thalamas for gross touch.

Vision is generated by light that hits the retina of the eye. Photoreceptors in the retina transduce the sensory stimulus of light into an electrical nerve signal that is sent to the visual cortex in the occipital lobe.

Visual signals leave the retinas through the optic nerves. Optic nerve fibers from the retinas' nasal halves cross to the opposite sides joining the fibers from the temporal halves of the opposite retinas to form the optic tracts.

The arrangements of the eyes' optics and the visual pathways mean vision from the left visual field is received by the right half of each retina, is processed by the right visual cortex, and vice versa.

The optic tract fibers reach the brain at the lateral geniculate nucleus , and travel through the optic radiation to reach the visual cortex. Hearing and balance are both generated in the inner ear.

Sound results in vibrations of the ossicles which continue finally to the hearing organ , and change in balance results in movement of liquids within the inner ear.

This creates a nerve signal that passes through the vestibulocochlear nerve. From here, it passes through to the cochlear nuclei , the superior olivary nucleus , the medial geniculate nucleus , and finally the auditory radiation to the auditory cortex.

The sense of smell is generated by receptor cells in the epithelium of the olfactory mucosa in the nasal cavity. This information passes via the olfactory nerve which goes into the skull through a relatively permeable part.

This nerve transmits to the neural circuitry of the olfactory bulb from where information is passed to the olfactory cortex.

Some taste information is also passed from the pharynx into this area via the vagus nerve. Information is then passed from here through the thalamus into the gustatory cortex.

Autonomic functions of the brain include the regulation, or rhythmic control of the heart rate and rate of breathing , and maintaining homeostasis.

Blood pressure and heart rate are influenced by the vasomotor centre of the medulla, which causes arteries and veins to be somewhat constricted at rest.

It does this by influencing the sympathetic and parasympathetic nervous systems via the vagus nerve. Information about the pressure changes in the carotid sinus comes from carotid bodies located near the carotid artery and this is passed via a nerve joining with the glossopharyngeal nerve.

This information travels up to the solitary nucleus in the medulla. Signals from here influence the vasomotor centre to adjust vein and artery constriction accordingly.

The brain controls the rate of breathing , mainly by respiratory centres in the medulla and pons.

This is a mixed nerve that carries sensory information back to the centres. There are four respiratory centres, three with a more clearly defined function, and an apneustic centre with a less clear function.

In the medulla a dorsal respiratory group causes the desire to breathe in and receives sensory information directly from the body.

Also in the medulla, the ventral respiratory group influences breathing out during exertion. In the pons the pneumotaxic centre influences the duration of each breath, [97] and the apneustic centre seems to have an influence on inhalation.

The respiratory centres directly senses blood carbon dioxide and pH. Information about blood oxygen , carbon dioxide and pH levels are also sensed on the walls of arteries in the peripheral chemoreceptors of the aortic and carotid bodies.

This information is passed via the vagus and glossopharyngeal nerves to the respiratory centres. High carbon dioxide, an acidic pH, or low oxygen stimulate the respiratory centres.

The hypothalamus in the diencephalon , is involved in regulating many functions of the body. Functions include neuroendocrine regulation, regulation of the circadian rhythm , control of the autonomic nervous system , and the regulation of fluid, and food intake.

The circadian rhythm is controlled by two main cell groups in the hypothalamus. The anterior hypothalamus includes the suprachiasmatic nucleus and the ventrolateral preoptic nucleus which through gene expression cycles, generates a roughly 24 hour circadian clock.

In the circadian day an ultradian rhythm takes control of the sleeping pattern. Sleep is an essential requirement for the body and brain and allows the closing down and resting of the body's systems.

There are also findings that suggest that the daily build-up of toxins in the brain are removed during sleep. Sleep necessarily reduces this use and gives time for the restoration of energy-giving ATP.

The effects of sleep deprivation show the absolute need for sleep. The lateral hypothalamus contains orexinergic neurons that control appetite and arousal through their projections to the ascending reticular activating system.

Through the autonomic projections, the hypothalamus is involved in regulating functions such as blood pressure, heart rate, breathing, sweating, and other homeostatic mechanisms.

The hypothalamus is influenced by the kidneys: when blood pressure falls, the renin released by the kidneys stimulates a need to drink. The hypothalamus also regulates food intake through autonomic signals, and hormone release by the digestive system.

While language functions were traditionally thought to be localized to Wernicke's area and Broca's area , [] it is now mostly accepted that a wider network of cortical regions contributes to language functions.

The study on how language is represented, processed, and acquired by the brain is called neurolinguistics , which is a large multidisciplinary field drawing from cognitive neuroscience , cognitive linguistics , and psycholinguistics.

The cerebrum has a contralateral organisation with each hemisphere of the brain interacting primarily with one half of the body: the left side of the brain interacts with the right side of the body, and vice versa.

The developmental cause for this is uncertain. Visual input follows a more complex rule: the optic nerves from the two eyes come together at a point called the optic chiasm , and half of the fibres from each nerve split off to join the other.

The left and right sides of the brain appear symmetrical, but they function asymmetrically. There are, however, several important exceptions, involving language and spatial cognition.

The left frontal lobe is dominant for language. If a key language area in the left hemisphere is damaged, it can leave the victim unable to speak or understand, [] whereas equivalent damage to the right hemisphere would cause only minor impairment to language skills.

A substantial part of current understanding of the interactions between the two hemispheres has come from the study of " split-brain patients"—people who underwent surgical transection of the corpus callosum in an attempt to reduce the severity of epileptic seizures.

Emotions are generally defined as two-step multicomponent processes involving elicitation , followed by psychological feelings, appraisal, expression, autonomic responses, and action tendencies.

The amygdala , orbitofrontal cortex , mid and anterior insula cortex and lateral prefrontal cortex , appeared to be involved in generating the emotions, while weaker evidence was found for the ventral tegmental area , ventral pallidum and nucleus accumbens in incentive salience.

The brain is responsible for cognition , [] [] which functions through numerous processes and executive functions.

The prefrontal cortex plays a significant role in mediating executive functions. Brain activity is made possible by the interconnections of neurons that are linked together to reach their targets.

Dendrites are often extensive branches that receive information in the form of signals from the axon terminals of other neurons. The signals received may cause the neuron to initiate an action potential an electrochemical signal or nerve impulse which is sent along its axon to the axon terminal, to connect with the dendrites or with the cell body of another neuron.

An action potential is initiated at the initial segment of an axon, which contains a specialized complex of proteins. The brain can also utilize lactate during exercise.

However, short-chain fatty acids e. The function of sleep is not fully understood; however, there is evidence that sleep enhances the clearance of metabolic waste products, some of which are potentially neurotoxic , from the brain and may also permit repair.

The brain is not fully understood, and research is ongoing. The boundaries between the specialties of neuroscience , neurology and other disciplines such as psychiatry have faded as they are all influenced by basic research in neuroscience.

Neuroscience research has expanded considerably in recent decades. The " Decade of the Brain ", an initiative of the United States Government in the s, is considered to have marked much of this increase in research, [] and was followed in by the BRAIN Initiative.

Information about the structure and function of the human brain comes from a variety of experimental methods, including animals and humans.

Information about brain trauma and stroke has provided information about the function of parts of the brain and the effects of brain damage.

Neuroimaging is used to visualise the brain and record brain activity. Electrophysiology is used to measure, record and monitor the electrical activity of the cortex.

Measurements may be of local field potentials of cortical areas, or of the activity of a single neuron. An electroencephalogram can record the electrical activity of the cortex using electrodes placed non-invasively on the scalp.

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These infrastructure development efforts will be aligned with the developments of the Human Brain Project and the Canadian Healthy Brains for Healthy Lives HBHL program to maximize compatibility with the international initiatives.

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